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It is [http://google.com Princess Peach's] birthday and as Mario, Luigi and their friends are celebrating, a huge cake slides in. Immediately, Bowser Jr. and the Koopalings pop out and throw the giant cake on top of Peach, trapping her inside. From there, the villains load her onto their ship while Mario, Luigi, Blue Toad and Yellow Toad follow closely. The Toads from the castle soon use a cannon to blast away the Propeller and Penguin Suits towards Mario and the others so they can have access to them.

When Mario and the gang finally get to the final castle, they find none other than Bowser himself. One of them has to jump on a ! Switch behind Bowser to defeat him, sending him falling into the lava below. After, Mario, Luigi, Yellow Toad, and Blue Toad celebrate their victory, they see Peach crying in a cage. However, she turns out to be Kamek in a pink version of his robe. Kamek then powers Bowser up, making him huge. After Bowser has been powered up, the four can do nothing but flee from the giant boss. Soon they find a dead end with a huge ! Switch. They push it, and Giant Bowser falls through the floor in final defeat as Princess Peach is released from her cage. As they venture out, they see Luigi, Blue Toad and Yellow Toad arrive in their hot air balloons. Luigi lets Mario and Peach into his balloon and is accidentally left behind. The two Toads then offer Luigi a ride in their balloon. As they sail off, Peach asks Mario if she told him about the Secret World.